The Reserve covers a large expanse of land running along the border between Zamora, Salamanca and Portugal. Both the Portuguese and Spanish areas are home to a large number of landscapes of immense orographic variety. The land has also been put to multiple uses, resulting in a wealth of elements and intriguing scenery.

Due to its vast size, it also boasts a large number of ecosystems and species reflecting an immense biological diversity. This area also features built heritage elements dating back to the days of the Romans and the Middle Ages. Examples include the remains of forts, castles and walled enclosures in towns such as San Felices de los Gallegos in the province of Salamanca.

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